| Name | Ownership | Shares Owned | sole voting power | shared voting power | Signature | Title | CIK |
|---|---|---|---|---|---|---|---|
| EcoR1 Capital, LLC | 12% | 7,803,202 | 0 | 7,803,202 | /s/ Oleg Nodelman | Manager | 0001587114 |
| Oleg Nodelman | 12% | 7,803,202 | 0 | 7,803,202 | /s/ Oleg Nodelman | Reporting Person | 0001454385 |
| EcoR1 Capital Fund Qualified, L.P. | 11% | 7,339,498 | 0 | 7,339,498 | /s/ Oleg Nodelman | Manager of the General Partner, EcoR1 Capital, LLC | 0001598292 |
| EcoR1 Capital Fund, L.P. | 0.7% | 463,704 | 0 | 463,704 | /s/ Oleg Nodelman | Manager of the General Partner, EcoR1 Capital, LLC | 0001567938 |