| Name | Ownership | Shares Owned | sole voting power | shared voting power | Signature | Title | CIK |
|---|---|---|---|---|---|---|---|
| Pfizer Inc. (IRS Identification No.: 13-5315170) | 3% | 157,646 | 26,537 | 131,109 | /s/ Madelyn D. Purcell | Madelyn D. Purcell/Assistant Secretary | |
| Pfizer Ventures (US) LLC (IRS Indentification No. 82-5524518 ) | 2.5% | 131,109 | 0 | 131,109 | /s/ Esther Louis | Esther Louis/ Secretary |